Each is also behind a locked door that requires you to have a Gardener’s Key from the Promenade of the Condemned. The three Moonflower Keys are hidden in secret rooms accessible through invisible passages, one in the Ramparts, one in the Graveyard, and one in the Forgotten Sepulcher. To get the Acceptance blueprint, you’ll need to find the three Moonflower Keys. The Acceptance mutation makes it so that you have to kill 50% fewer enemies to remove a curse than you otherwise would.

You can get it by defeating the Elite Undead Archer in the Ramparts. You can get it by defeating the Hand of the King, the last of the game’s current bosses, in the Throne Room.Ĭustomization Rune (added in version 1.2) – With the Customization Rune, you’ll unlock Custom Mode, which is accessible from the main menu and lets you create custom runs. Homunculus Rune – The Homunculus Rune gives you the ability to move your character’s head separately from their body, detaching to travel a set distance away (if you take the head too far, it will automatically return), where it can move turrets, grab loot, grab blueprints, stun enemies, and access areas too small for your body to enter. You can get it by defeating the Elite Caster in the Slumbering Sanctuary, in a room where you will need to use your newfound wall jump to leave. Spider Rune – The Spider Rune lets you wall jump and climb walls, granting you access to the Graveyard, the Prison Depths, and the Slumbering Sanctuary when entered via Black Bridge. You can get it by defeating the Elite Slasher in the Ossuary. You can get it by defeating The Concierge, one of the game’s bosses, found at Black Bridge.īelier’s Rune/Ram Rune (renamed) – The Ram Rune increases the power and range of your Dive attack and lets you use it to break through fragile areas, granting you access to the Ancient Sewers.

You can get it by defeating the Elite Slasher in the Toxic Sewers.Ĭhallenger Rune – The Challenger Rune opens the door in the Prisoners’ Quarters that lets you play the Daily Run competitive multiplayer mode. Teleportation Rune – With the Teleportation Rune, you can teleport between certain special coffins, granting you access to the Ossuary and the Forgotten Sepulcher. You can get it by defeating the Elite Undead Archer in the Promenade of the Condemned. Vine Rune – The Vine Rune gives you the ability to make vines grow from certain green plant clumps in the game, granting you access to the Toxic Sewers and the Ramparts. There are seven runes in the game, each with a different ability.

To get a rune, you will need to go to a specific location and fight the Elite enemy that spawns when you approach the special obelisk in the area, or by fighting a boss. These upgrades give you new abilities that let you access areas you otherwise couldn’t reach. Runes are permanent upgrades attached to your save file.

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